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Earthworms ingest 10.3% of annual average broad-leafed litter in Liangshui national nature reserve. 在涼水國家級自然保護區，蚯蚓對濶葉凋落葉的分解佔年平均凋落葉量的10.3%左右。
Thispaper proposed the idea that broad - leafed forest and mixed forest could be bred by artificalfacilitation on the same condition. 同時提出了在類似條件下通過人工促進天然更新培育濶葉林和針濶混交林的設想。